Cordillera Oriental montane forests

Achiote, also sold as annatto or lipstick tree, is a fast-growing evergreen shrub or small tree native from Mexico through Central America to Colombia, Venezuela, the Guianas, Ecuador, Peru, Bolivia, Brazil, Paraguay, and northern Argentina, and planted across the tropics for its seeds. Clusters of pink or white flowers about 2 inches across, like single wild roses, open at the branch tips and are followed by clusters of red to red-brown capsules covered in soft bristles, which split open to show seeds coated in a waxy red-orange layer. That coating is annatto: the coloring and mild seasoning behind achiote paste and sazón, the yellow-orange of many cheeses, butters, and margarines, and the red body paint and lipstick that gave the plant its English name. In a garden it usually stays 6 to 12 feet tall and wide as a specimen or informal hedge, though as a tree it can reach 20 to 30 feet. It is frost-tender and needs a frost-free climate: it is hardy outdoors in USDA zones 10a-12b and grown elsewhere in a large container or conservatory that comes indoors before frost. Outside its native range it has naturalized on many Pacific islands, sparingly so in Hawaii, and in the Galapagos, where the Charles Darwin Foundation rates it potentially invasive.

A bold, clump-forming evergreen perennial from South Africa, grown for big rounded umbels of trumpet-shaped blue (or white) flowers held on tall bare stalks above arching, strap-shaped leaves in mid-to-late summer. It is widely sold as "lily of the Nile," but that is a misnomer - the plant is South African (the Cape provinces and KwaZulu-Natal), not from the Nile. Spectacular and easy in warm climates, this evergreen Agapanthus is frost-tender, so in cold-winter areas it is grown in a container and overwintered under cover. The RHS has given several Agapanthus praecox forms its Award of Garden Merit and rates this evergreen species half-hardy (H3 - needs winter protection).

One of the world's most spectacular flowering trees and, in the same breath, one of its most dangerous invaders. Spathodea campanulata is a large, fast-growing tropical tree native to the moist forests of West and Central Africa (GBIF), carrying a dense crown of glossy compound leaves topped by showy, upturned clusters of big, frilly, scarlet-to-orange, tulip-shaped flowers (the boat-shaped buds hold water). It is glorious - and it is a serious invasive. The African tulip tree is listed among the IUCN "100 of the world's worst invasive species"; outside its native range it has overrun wet-tropical forests across the Pacific, Hawaii, the Caribbean, and parts of Asia and Australia, and its flowers can be lethal to native bees. Plant it ONLY where it is genuinely native or where it cannot escape, and first check that it is not prohibited locally - do not plant it in any tropical region where it can naturalise. It is frost-tender (USDA 10a-11), fast, and wants full sun and moisture; in cold-winter climates it is grown only as a tender container specimen kept frost-free. It is an ornamental, not a food plant.

Aji pepper is the South American chili species behind aji amarillo, Lemon Drop, Bishop's crown and Brazil's dedo-de-moça - one of the five domesticated chili peppers, and a different plant from the familiar garden pepper (Capsicum annuum) and the habanero (Capsicum chinense). It is a perennial shrub native from Colombia, Ecuador and Peru through Bolivia, Paraguay, Uruguay, northern Argentina and central and southern Brazil, and where it can live for years it grows taller than most garden peppers, up to about 6.6 feet. Its white to cream flowers are typically marked with green or gold, and its fruits, which usually hang down in cultivated forms, come in an unusually wide range of shapes, from long tapered pods to the crown-shaped Bishop's crown. Heat runs from nearly heatless types to hot ones; aji amarillo is rated around 30,000-50,000 Scoville units. It is frost tender, so outside nearly frost-free climates it is grown as a warm-weather annual.

Hippeastrum x hybridum is a complex garden hybrid derived from six or more wild Hippeastrum species native to tropical and subtropical South America, developed through intensive breeding since the late 18th century and now the dominant "amaryllis" sold worldwide as a holiday bulb. It produces spectacular trumpet-shaped flowers - 5 to 8 inches across - on stout leafless scapes in red, pink, white, orange, salmon, or bicolour, making it one of the most dramatic winter-flowering houseplants available. The honest catch is toxicity: the entire plant, especially the bulb, contains lycorine and related alkaloids at concentrations that cause vomiting, salivation, tremors, and hypotension in pets and children, so it must be placed out of reach and handled with care.

Angel's trumpet (Brugmansia suaveolens) is a large, soft-wooded shrub or small tree hung with enormous pendulous trumpet flowers - up to 12 inches long, white aging to soft blush, and intensely fragrant in the evening and at night. Native to the Atlantic Forest of south-eastern Brazil (POWO, Kew; Flora e Funga do Brasil), it is now classified Extinct in the Wild by the IUCN and survives only in cultivation, which makes a familiar garden plant quietly remarkable. It grows fast and thirsty in warmth (USDA zones 9b-11b) and is frost-tender, so in cold-winter areas it is grown in a large container and overwintered under cover. The load-bearing warning is its toxicity: EVERY PART IS HIGHLY TOXIC, loaded with tropane alkaloids (scopolamine, atropine, hyoscyamine), and poisonings can be severe or fatal - never plant it where children or pets could reach it.

A tender perennial from Madagascar grown across temperate North America as a heat-loving summer annual - a mounding 6-18 inch plant in the dogbane family covered in flat five-lobed phlox-like flowers from June to frost. The species blooms rosy-pink to red with a darker mauve throat, and it shrugs off the hot, humid weather that wilts most bedding plants. Every part of the plant is poisonous: it is the natural source of the vinca alkaloids used in chemotherapy.

A fast cool-season annual of the mustard family grown for its peppery, mustard-like salad greens - irregular, pinnately-lobed basal leaves in a low rosette, each with 4 to 10 small lateral lobes and a large terminal lobe (Missouri Botanical Garden). First cultivated by the ancient Greeks and Romans and still widely grown across Europe, it is best grown in the cooler spring and fall months rather than summer heat; leaves are harvested young and tender before they turn strong and bitter. Pale-yellow four-petalled flowers with dark brown or purple veins appear in corymbs if plants are left to bloom.

A frost-tender broadleaf evergreen tree of the laurel family, native to Mexico and Central America and grown across the tropics and subtropics for its buttery, pear-shaped fruit. Glossy dark-green elliptic leaves 4 to 8 inches long clothe a tree that reaches 30 to 60 feet, hung with greenish-yellow flower panicles that give way to large single-seeded berries. Hardy only in USDA zones 10-12 - north of that it is an indoor curiosity easily sprouted from a pit, but one that rarely fruits.

A giant herbaceous perennial from Southeast Asia and the principal wild ancestor of most cultivated dessert bananas. What looks like a trunk is a 'pseudostem' - tightly rolled leaf sheaths - topped by a fountain of huge, paddle-shaped leaves that can run 6 to 10 feet long, giving an instant tropical effect. In frost-free climates (USDA zones 10a-11b) an established clump produces a drooping flower spike and a hanging bunch of edible fruit, then that pseudostem dies and is replaced by a sucker from the base. It is frost-tender: everywhere colder it is grown as a bold container or greenhouse foliage plant that is overwintered indoors and rarely, if ever, fruits.

Gerbera jamesonii, the Barberton daisy (also Transvaal daisy), is a tufted evergreen perennial herb in the daisy family (Asteraceae) native to the summer-rainfall grasslands and rocky woodland of north-eastern South Africa and Eswatini. It forms a basal rosette of lobed leaves from which leafless flowering scapes rise, each topped by a single large daisy-style flowerhead in orange-red, yellow, pink, or white. It is the wild ancestor of the thousands of florist gerbera cultivars and earns its place as a long-blooming focal point in borders and patio containers, attractive to bees and other insects. The load-bearing caution is frost-tenderness: RHS rates it H1C, meaning it survives outdoors only in summer or the very mildest, frost-free spots and must be overwintered under glass elsewhere (roughly USDA 9-11). It is non-toxic, with no reported poisoning hazard to people or pets, making it a safe choice where toxicity is a concern.

A woody, deciduous flowering shrub in the Hydrangeaceae, native to Japan, China, Korea, and Southeast Asia and long grown as the classic "hortensia" or French hydrangea. NC State Extension describes a rounded shrub 3 to 6 feet tall and wide with large opposite, simple, toothed leaves (4 to 8 inches long) and big rounded mop-head or flat lacecap flower clusters in late spring and summer in white, pink, blue, or purple. Famously, flower color tracks soil chemistry - acidic soils push the blooms blue and alkaline soils turn them pink. It wants protection from hot afternoon sun and steady moisture, making it a mainstay of shaded foundation plantings and woodland borders.

A tall, summer-growing cormous perennial from across sub-Saharan Africa, grown for arching spikes of large, hooded, orange-to-red-and-yellow flowers in summer over fans of sword-shaped leaves. This is the widespread wild African gladiolus and a key ancestor of the big garden gladioli. Unlike the winter-growing Cape bulbs, it is a SUMMER-GROWING corm: plant in spring, grow and flower through summer, then it dies back to a dormant corm for winter. It wants full sun, well-drained soil, and summer moisture, and is one of the hardier species gladioli - surviving to about USDA zone 7 with a deep winter mulch or by lifting the corms in cold-winter areas. The corms are toxic and irritant if eaten and have been used in traditional medicine, never as food. In the wild the hooded flowers are pollinated by sunbirds and long-tongued insects.

American beautyberry (Callicarpa americana) is a loose, open, arching deciduous shrub of the southeastern and south-central United States, grown almost entirely for one spectacular autumn trick. Through summer it is a quiet green background plant, carrying small clusters of pink-to-lilac flowers tucked into the leaf axils. Then, from late summer into fall, those clusters ripen into tight whorls of vivid magenta-purple berries that wrap right around the stems at every leaf joint, so the arching branches look strung with beads. Songbirds strip them through fall and early winter. It is an easy, fast-growing, forgiving native that asks for little: full sun to part shade, ordinary well-drained soil, and room to arch. Site it where the fall berry show can be seen, because in leaf and flower it is plain, and in fruit it is one of the most distinctive shrubs you can grow.

Dierama pulcherrimum, angel's fishing rod, is an arching, more-or-less evergreen cormous perennial in the iris family (Iridaceae), endemic to the open summer-rainfall grasslands of the Eastern Cape of South Africa. It forms grassy tufts of long, narrow leaves from which tall, wiry, arching flower stems rise to about 4.9 feet (occasionally to 5.9 feet), each hung with pendulous, bell-shaped rosy-pink to purple flowers in summer - the namesake "fishing rod" silhouette over a pond or in a gravel garden. It is the load-bearing caution to note that this is only a borderline-hardy plant: RHS rates it H4 (hardy to roughly 14 to 23°F / USDA ~8b-9), so in colder zones it needs a sheltered, sunny, sharply drained spot, a winter mulch, or container/glasshouse protection, and it resents disturbance once established. It is not recorded as toxic - though it is grown purely as an ornamental and is not a food plant, so treat it as inedible. It is well-behaved (Least Concern in the wild, no recorded weediness) and suits borders, pond margins, and containers in mild-temperate gardens.

Anise-scented sage is a tall, upright perennial in the mint family (Lamiaceae), native to Brazil, Paraguay, Uruguay, and Argentina. NC State Extension and Missouri Botanical Garden PlantFinder describe a bushy, square-stemmed plant 2 to 5 feet tall and wide that carries spikes of deep blue, two-lipped flowers with purple-blue calyxes from July into fall; NC State calls it a reliable nectar source for hummingbirds and butterflies, and notes that carpenter bees rob its nectar by biting a hole in the floral tube. The 'anise-scented' name is only partly earned: Wikipedia describes 'an anise scent when crushed,' but Missouri Botanical Garden PlantFinder says the bruised foliage has a very mild aroma with 'very little if any anise scent.' It spreads by tuberous roots that let it reproduce vegetatively (NC State Extension), so it fills a space rather than staying in a tidy clump. It is winter hardy only to about USDA zone 8a: farther north, grow it as an annual or overwinter cuttings indoors, though NC State notes the 'Black and Blue' cultivar is hardier and will survive milder zone 7 winters if well mulched. Neither NC State nor Missouri Botanical Garden documents an edible use or a toxicity hazard. NC State records, as ethnobotanical history, that the Guarani people of Brazil used it as a sedative; that is not a recommendation to take it, so treat it as a non-food ornamental. It has also naturalized outside South America (Wikipedia records it in New Zealand and Chile), so where winters are mild enough for it to persist outdoors, check your local weed or invasive-plant list before planting it.

A deciduous Eastern-Asian fruit tree with a rounded, spreading crown that the Missouri Botanical Garden lists at 20 to 30 feet tall and wide. Oval leaves emerge yellowish-green, mature to glossy green, and turn gold to red in fall; fragrant but insignificant late-spring flowers give way to showy orange persimmons (3 to 4 inches) that ripen in late fall and may persist on bare branches into winter. Winter hardy to USDA Zones 7-10 and drought tolerant once established.

The Mediterranean evergreen whose leathery, glossy dark-green leaves are the bay leaf of the kitchen. Missouri Botanical Garden PlantFinder describes it as a pyramidal, aromatic evergreen tree or large shrub that can reach 60 feet but is usually seen at 10 to 30 feet and is often pruned to 8 feet or less for garden use. Trees are dioecious: small yellowish-green spring flowers on female plants, if pollinated, give way to single-seeded purple-black berries. Winter hardy only to USDA Zone 8, so it is grown as a clipped container houseplant farther north.

Bear's breeches is a bold Mediterranean perennial whose dramatically lobed, glossy leaves inspired the Corinthian column capital of classical architecture. Its wild native range spans the central and eastern Mediterranean - Italy, Sicily, Sardinia, the Balkans, Greece, the Aegean, Anatolia and the Levant, plus NW Africa (Morocco, Algeria, Tunisia); it is naturalised (not native) further west into Iberia. In gardens it serves as an architectural statement plant, producing tall white-and-purple flower spikes in summer on established clumps. The honest catch is its root system: fleshy, deep-running tuberous roots regenerate readily from the smallest fragment, making Acanthus mollis genuinely difficult to eradicate once established and a declared invasive in Australia and New Zealand, so site selection is a permanent decision.

The classic German or common-flag iris - the presumed parent of most modern bearded-iris cultivars, probably native to southern Europe and the Mediterranean and naturalized widely. Each stalk carries up to six large, usually fragrant flowers in spring: three erect lilac standards above three purple falls marked with brown veins, white bases, and the signature yellow "beard." It has no bulb, spreading instead by creeping rhizomes that form large clumps, with sword-shaped basal foliage to about two feet.

Boxelder (Acer negundo) is the maple that does not look like a maple. Instead of a single lobed leaf it carries compound leaves of three to five toothed leaflets, close enough to poison ivy that people regularly misidentify young shoots, and it is one of the very few dioecious maples - male and female flowers on separate trees, so only the females carry the familiar paired samaras. It is native across an extraordinary sweep of North America, from Maine to Manitoba and south to Florida, Texas and central Mexico, and it is genuinely at home on riverbanks and floodplains where its speed and flood tolerance are assets. In a garden it is usually a volunteer rather than a choice. It grows fast, breaks in storms, seeds everywhere, and hosts the boxelder bug, which overwinters on house walls in numbers that make it a household nuisance rather than a garden one. The case for it is narrow but real: it establishes on ground where better trees fail, in cold northern climates and on raw disturbed sites, and it is a documented larval host for cecropia and a valued early honey-bee forage.

A bold, clumping rhizomatous perennial grown for its elegant, pure-white funnel-shaped flowers - actually a curved spathe wrapped around a yellow finger-like spadix - held on tall stems above large, glossy, arrow-shaped deep-green leaves from late winter into summer. Native to the wetlands and seeps of southern Africa, Zantedeschia aethiopica is the classic florist's calla and a striking water-margin and container plant, thriving in consistently moist to wet rich soil in full sun to part shade. It is tender (hardy in zones 8a-10b), where it overwinters in the ground and can be near-evergreen; colder gardeners grow it from lifted rhizomes or under heavy mulch. Two cautions are load-bearing: every part is toxic if eaten (calcium-oxalate raphides), and in mild, wet climates it can escape and naturalize into an invasive weed.

A tall, statuesque, winter-growing cormous perennial from the winter-rainfall fynbos of South Africa's Western Cape, grown for its bold fans of upright sword leaves and branched spikes of pink (to white) tubular flowers in late spring. Known as the Cape bugle-lily, it is a striking fynbos geophyte that flowers profusely the season after a fire. Honesty (load-bearing): it is a TALL plant, 4 to 6 feet in flower, and it is WINTER-GROWING and SUMMER-DORMANT - it grows through autumn and winter, flowers in late spring, then dies back and rests dry through summer, so you water it from autumn to spring and keep it dry and at rest in summer; watering it through a hot summer rots the corm. It is FROST-TENDER to borderline (RHS rates it about H3-H4): in cold-winter areas grow it in a warm, sheltered, sharply-drained spot or a large pot, or lift and store the dormant corms. It wants FULL SUN and well-drained soil and the room its height needs. In the wild it is FIRE-ADAPTED, flowering en masse the season after a fynbos fire. The corms are inedible. It is bird-pollinated, worked by sunbirds (and butterflies) along with long-tongued flies - described in the prose, since it is grown as an ornamental rather than for any crop.

Cider gum is a fast-growing evergreen tree from the woodlands of Tasmania's central plateau and one of the most cold-hardy eucalypts: Trees and Shrubs Online calls it the hardiest of the tree-form eucalypts planted widely enough to judge, and the RHS rates it H5, hardy in most of the UK even in severe winters. Plants For A Future gives it long spells down to 7°F and short ones down to -0°F. Left unpruned it grows into a tree of 40 to 100 feet with peeling cream and brown bark and grey-green, sickle-shaped adult leaves, but many gardeners cut it back hard every spring to keep it in the rounded, glaucous-blue juvenile foliage it is best known for. Small white flowers in clusters of three appear on trees allowed to mature, usually in summer or autumn in British gardens. The name refers to its sweet sap, which the Aboriginal people of Tasmania fermented into a drink called way-a-linah. The leaves hold eucalyptus oil, which is poisonous if swallowed (Plants For A Future reports deaths after ingestion of 4-24 ml of eucalyptus oils), so keep foliage, prunings and any oil away from children and pets. Outside its range it has naturalized in woodland and on roadsides in south-eastern England, where it occasionally regenerates from seed, while in the wild it is assessed as Endangered on the IUCN Red List.

Cacao is the chocolate tree - a small evergreen understory tree of the lowland tropical rainforest, famous for being cauliflorous: its tiny pink-and-white flowers, and then the large ridged football-shaped pods, are borne directly on the trunk and oldest branches rather than out among the leaves. Reaching roughly 15 to 25 feet, it carries glossy oblong leaves that flush bronze when young. Each pod holds 30-40 seeds embedded in sweet white pulp; those seeds become chocolate only after fermenting, drying, and roasting (the cacau of Brazil). It is genuinely demanding - a true tropical, needing deep humid warmth and overhead shade and failing below about 50°F - so outside the wet tropics it is a greenhouse or conservatory plant only. Its flowers are pollinated not by bees but by tiny midges.

The classic florists' gloxinia, a tuberous tropical perennial of the Gesneriaceae native to the Atlantic Forest of south-eastern Brazil (POWO, Kew; recorded as Brazilian-native in Flora e Funga do Brasil). It forms a low, neat rosette of soft, velvety leaves above which sit large, showy, bell-shaped flowers in rich violet, red, pink or white, often finished with a contrasting picotee edge. POWO and Flora e Funga do Brasil record Sinningia speciosa as the wild parent of the cultivated florist gloxinia, and that ornamental heritage is exactly how it is grown almost everywhere - as a tender pot or houseplant prized for its plush leaves and velvety trumpet-and-bell flowers. HONESTY: it is very frost-tender, hardy in the ground only in roughly USDA zone 11 and above, and it has a real dormant season - after flowering it dies back to a resting tuber before regrowing, so it is not an evergreen plant but a cyclical one. It is grown purely as an ornamental: no part is eaten. The flowers are insect-pollinated and, in the plant's native Brazilian range, also visited by hummingbirds.

Snowflower (also called peace lily, a name used across its genus) is a wild tropical aroid that Kew's WCVP places native to Colombia, Ecuador, Peru, and Venezuela. It is a clump-forming evergreen perennial whose oval to lance-shaped leaves are 5 to 8 inches long, at the small end of a genus whose leaves run 5 to 26 inches, and its spathe is greenish- or yellowish-white rather than pure white, 4 to 8 inches long around the flowering spadix. Many peace lilies in the houseplant trade are cultivars and hybrids, so a plant sold simply as a peace lily is not necessarily this species. Like the rest of its genus it contains calcium-oxalate crystals, a genus-level finding that makes it mildly toxic to people and pets if chewed, and it is frost-tender: hardy outdoors only in frost-free USDA zones 11-12 and grown as a houseplant everywhere else.

Tree thorn-apple (Brugmansia arborea), also known as angel's trumpet, horn of plenty or maikoa (RHS), is an evergreen to semi-evergreen shrub or small tree of the Andes, first described by Linnaeus as Datura arborea. Kew records it as native to Bolivia, northern Chile, Ecuador and Peru, and Wikipedia describes it from drier valleys at 6,562 to 9,843 feet that often see light frost. In gardens RHS gives it an ultimate size of 8.2 to 13.1 feet tall and 4.9 to 8.2 feet wide; in its home valleys it can reach 23 feet. Its strongly fragrant, nodding white to cream trumpets are only 5 to 7 inches long, the shortest in the genus, and each sits in a very long green calyx that reaches almost to the mouth of the flower - the quickest way to tell it from the many other white angel's trumpets sold under its name. It flowers almost continuously in small numbers rather than in big flushes. The load-bearing warning is its toxicity: RHS lists it as TOXIC if eaten, a skin irritant and toxic to pets, and every part of a Brugmansia is potentially poisonous, carrying tropane alkaloids (scopolamine, hyoscyamine, atropine), with the seeds and leaves especially dangerous - never plant it where children or pets could reach it. The IUCN Red List classes it as Extinct in the Wild, yet it is listed as invasive in New Caledonia. Wikipedia calls it the hardiest Brugmansia to both cold and drought, but RHS rates it H1c, wanting a winter minimum of about 41°F, so Plotwright publishes USDA zone 11 only and treats it as a container plant to bring under cover anywhere colder.