Helanshan montane conifer forests

A tall, bold warm-season annual from Mexico and Guatemala (the "African" name is a misnomer of its European garden history) grown for large, fully double, pompon-like flowerheads in saturated yellow, gold, and orange over strongly aromatic, finely divided foliage. Plants reach 1 to 4 feet and bloom from early summer to frost in full sun. The petals are edible and used as a culinary garnish and natural dye, and the flowers are the iconic "flor de muerto" of Mexican Day of the Dead. Despite the wide listed zone range it is frost-tender and grown for a single warm season.

A native eastern + central North American thicket-forming shrub producing arching thorny canes + clusters of large sweet black berries in mid-to-late summer. Among the most important wildlife fruit producers in eastern forests - birds, mammals, + insects all depend on the fruit. Like raspberry, biennial-caned (primocane year 1, fruits in year 2 as floricane, then dies back). Spreads via root suckers + tip-rooting cane tips; manage with annual pruning.

A dense, conical-to-narrow-pyramidal evergreen tree native to eastern and central North America, prized as a screening and foundation conifer. Flat, fan-like sprays of scale-like, aromatic yellow-green foliage clothe the tree from the ground up, and red-brown bark exfoliates on mature trunks. Wild trees can reach 40 to 60 feet but cultivated plants typically stay near 20 to 30 feet; small urn-shaped cones and dense evergreen cover make it valuable food and shelter for birds.

A medium-to-large native shade tree of central and eastern North America, reaching 50 to 80 feet with an ovate-rounded crown and large, asymmetric heart-shaped leaves. In June it carries pale-yellow, intensely fragrant flowers on pendulous cymes - each cluster hung from a distinctive strap-like leafy bract - that ripen into pea-sized nutlets. The fragrant June bloom is a premier nectar source: Missouri Botanical Garden lists it as attracting bees and butterflies, and the Lady Bird Johnson Wildflower Center flags it as having special value to both native and honey bees.

A fast, suckering native shrub of streambanks and moist thickets across eastern North America, grown for huge flat-topped cymes of tiny lemon-scented white flowers in early summer and the clusters of dark elderberry drupes that follow. Spreads by root suckers into naturalized colonies 5 to 12 feet tall and wide; the flowers feed butterflies and the showy fruit feeds birds. The raw berries are not eaten fresh - they are cooked into jelly, pie, and wine.

The American elm is the great vase-shaped shade tree that once arched over Main Streets and town commons across eastern North America - a fast, extremely cold-hardy deciduous tree of 60 to 80 feet whose upright trunk divides into a fountain of high, spreading limbs that meet overhead to form a living cathedral ceiling. That iconic form, and the species' tolerance of wet soil and tough urban conditions, made it the default American street tree for a century. Then Dutch elm disease (DED) - an introduced fungal disease carried by elm bark beetles - swept through in the 20th century and killed the vast majority of mature street and shade elms across the continent. The honest reality for a gardener today is blunt: do not plant the unselected wild species expecting it to survive. If you want the American-elm form, plant a DED-tolerant cultivar bred and selected for resistance - 'Princeton', 'Valley Forge', 'New Harmony', or 'Jefferson' - and say so plainly. Where it does grow, it is fast, hardy to USDA zone 3, and remarkably forgiving of wet ground and city stress.

American germander, also called wood sage, is a widespread North American native perennial in the mint family that runs steadily underground on creeping rhizomes. From early to midsummer it sends up erect, softly hairy stems topped with one-sided spikes of pale pink-to-lavender flowers, each with the distinctive deeply lobed lower lip that gives the germanders their look and makes a generous landing platform for bees. It is a plant of moist open ground - wet meadows, streambanks, ditches, and the edges of thickets - across most of the contiguous United States into southern Canada, which tells you exactly what it wants: sun and a soil that does not dry out. The honest caveat is its vigor: those same rhizomes that fill a bank or a rain garden so readily will also colonize a tidy perennial border and crowd politer neighbors. Site it where it can run, or give it a root barrier, and it rewards you with a long, dependable bee-friendly bloom rather than a maintenance fight.

A rounded, multi-stemmed deciduous shrub native across eastern and central North America, grown for its edible nuts and its season-opening catkins. Showy 2-3 inch yellowish-brown male catkins dangle from bare branches in early spring before the ovate, double-toothed leaves emerge; small egg-shaped edible nuts ripen inside leafy husks by mid- to late summer. Easygoing in average soil and tolerant of clay and black walnut, it suckers into thickets that screen and shelter wildlife.

A small-to-medium understory tree of dry, rocky eastern-North-American woods, named for its drooping clusters of papery, sac-like seed pods that resemble the fruit of hops. The birch-like, sharply-serrated leaves turn an undistinguished yellow in fall, and reddish-brown male catkins persist on the bare branches through winter. Also called ironwood for its extremely hard, dense wood; tough, low-maintenance, and drought-tolerant once established.

A tough, medium-sized native tree of the eastern and midwestern United States, grown as much for its showy edible orange fruit as for its distinctive thick, dark gray bark broken into rectangular blocks. Small urn-shaped white-to-greenish-yellow flowers open in May and June, and the sweet fruit ripens after frost. Largely dioecious - a female tree needs a male pollinizer nearby to set fruit - and notably drought- and walnut-tolerant once established.

A small native deciduous tree (or thicket-forming, suckering shrub) of eastern and central North America, grown for clouds of fragrant white 5-petaled flowers that open in March before the leaves and for the edible red plums that follow in early summer. It forms a broad, spreading crown with attractive dark reddish-brown twigs that sometimes carry thorny lateral branchlets. A documented larval host for swallowtails and other butterflies, with flowers of special value to native, bumble, and honey bees.

A native bramble (cane) producing red aromatic edible fruit in summer or fall (depending on summer-bearing vs everbearing cultivar). Self-pollinating; spreads vigorously by root suckers + tip-rooting canes. NC State documents extensive Lepidoptera + small mammal + bird wildlife value alongside the edible fruit role. Site where the spreading habit is welcome - naturalized colonies form in sun-exposed open ground.

A massive native deciduous canopy tree of eastern North American floodplain forests producing distinctive mottled white-tan-gray exfoliating bark (the design-defining trait - sycamore bark looks like military camouflage), large palmate maple-like leaves, and persistent spherical seed balls. Among the largest deciduous trees in eastern North America - old-growth specimens exceed 150 feet tall + 10 feet trunk diameter. Site only where massive scale is acceptable.

Amur maple (Acer ginnala, accepted by Kew's Plants of the World Online as Acer tataricum subsp. ginnala) is a small, fast-growing deciduous tree or large multi-stemmed shrub from northeastern Asia - the Russian Far East, northeastern China, and Korea. At 15 to 20 feet it fits where a full-size maple cannot, and it earns that place with three seasons of interest: fragrant greenish-yellow flowers in spring, which is an unusual scent for a maple and an early nectar source for bees; small glossy three-lobed leaves through summer; and reliable, often brilliant red-to-orange fall color that arrives early. It is also one of the hardiest maples, tolerant of cold, wind, dry soil, and a wide range of pH. The honest catch, and it is a serious one for North American gardeners, is that it self-sows prolifically and has escaped cultivation to become invasive across parts of the US Midwest and Northeast, where its shade-tolerant seedlings crowd native shrubs and understory trees in open woods, savannas, and prairies. Several states list it as invasive or a noxious weed. Check your local invasive-species list before planting, and be prepared to remove seedlings if you do.

An upright, clump-forming perennial of the mint family native to the upper Midwest, Great Plains, and into central Canada, named for its anise-scented foliage. From June through September it carries dense terminal spikes of lavender-to-purple two-lipped flowers above square stems and opposite, toothed leaves. The Lady Bird Johnson Wildflower Center flags it as a nectar source with special value to native bees, bumble bees, and honey bees, and it also draws butterflies and hummingbirds.

A native eastern-US deciduous shrub - 'Annabelle' is a sterile-flowered cultivar of smooth hydrangea - with very large white snowball blooms in summer. Blooms on new wood so spring frost cannot destroy the flower display, and serves as the larval host for the hydrangea sphinx moth.

A native central + eastern US perennial with intensely aromatic foliage when crushed and dense clouds of small blue-purple flowers in late fall - often the latest-blooming aster in the eastern flora. Drought + clay tolerant; among the toughest native fall pollinator plants.

A native eastern + central North American multi-stemmed deciduous shrub with dentate (toothed) foliage, white spring flower clusters, blue-black drupes, and reliable fall color. Especially valued for wildlife - among the most-cited native shrubs for fall-migration bird forage.

A fast cool-season annual of the mustard family grown for its peppery, mustard-like salad greens - irregular, pinnately-lobed basal leaves in a low rosette, each with 4 to 10 small lateral lobes and a large terminal lobe (Missouri Botanical Garden). First cultivated by the ancient Greeks and Romans and still widely grown across Europe, it is best grown in the cooler spring and fall months rather than summer heat; leaves are harvested young and tender before they turn strong and bitter. Pale-yellow four-petalled flowers with dark brown or purple veins appear in corymbs if plants are left to bloom.

Asiatic hybrids are the easiest lilies to grow and among the first to bloom - rigid, unbranched 3-4 foot stems carry large, mostly upward- and outward-facing flowers 4 to 6 inches wide in nearly every color but blue, often with dark basal spotting. The flowers are showy and good for cutting but, unlike most other lily groups, usually have little or no fragrance. Every part of the plant is dangerously toxic to cats.

A long-lived herbaceous perennial vegetable grown for the tender young spears harvested in April and May before they unfurl. Native to Europe and temperate Asia, it grows from a crown that takes 2-3 years to come into production but then yields for fifteen years or more. Spears left uncut grow into airy 3-4 foot summer ferns; the plants are dioecious, and female plants ripen ornamental red berries in late summer.

Aubrieta (Aubrieta deltoidea) is a low, mat-forming evergreen perennial in the mustard family (Brassicaceae), native to the rocky hillsides of southeastern Europe - primarily Greece, the Aegean Islands, Crete, and adjacent Mediterranean coasts. It is one of the most reliable spring-flowering ground covers for sunny, well-drained spots: cascading sheets of violet to deep pink four-petalled blooms from March through May, attractive to bees and bee flies. The honest catch is that without a hard cut-back immediately after flowering, plants become woody and bare in the centre within two or three years, collapsing from a tight carpet into a tired, gappy mat.

Austrian pine (Pinus nigra), also called European black pine, is a dense, dark, 40-to-60-foot conifer from central and southern Europe into western Asia, with stiff dark green needles three to six inches long in bundles of two and bark that darkens to nearly black on mature trunks. For most of the twentieth century it was the default tough conifer of North American landscapes: it takes clay, drought, salt and city air, and Missouri Botanical Garden still lists urban tolerance among its attributes. That reputation is now out of date, and the reason is a disease. Sphaeropsis - Diplodia - tip blight kills the new shoots from the bottom of the tree upward, hollowing out the crown over years, and it is severe enough that Missouri Botanical Garden explicitly no longer recommends planting Austrian pine in the St. Louis area or the wider Midwest. Dothistroma needle blight and pine wilt nematode compound it. It remains a legitimate choice on the drier, cooler, less humid end of its range; specifying it into a humid continental summer is planting a tree with a known ending.

Colchicum autumnale is a corm-forming herbaceous perennial native to lowland grassy meadows across much of Europe, from Portugal and Great Britain east to Ukraine. In autumn it sends up naked goblet-shaped flowers of lilac-pink directly from the bare soil - leaves and seedpods follow in spring and die back by early summer. The honest catch is its extreme toxicity: every part of the plant contains colchicine, a compound lethal to humans and animals, and the broad strap-like spring leaves are routinely mistaken for edible wild garlic - a potentially fatal confusion. Despite the common name, it is not a true crocus (Crocus, Iridaceae) but a member of the Colchicaceae.

Balsam fir (Abies balsamea) is THE Christmas tree of the northeast and, for most gardens, a tree to admire rather than plant. Missouri Botanical Garden is unambiguous about why: the species is native to cool climates and is not recommended for planting in the hot and humid summer conditions south of USDA Zone 5, it is generally intolerant of urban pollution, and MBG explicitly does not recommend it for its own St. Louis climate. Wikipedia puts the same limit the other way round - it wants a mean annual temperature near 40°F with consistent moisture, and low moisture combined with high temperature is what stops the range. This is a boreal tree. It runs from Labrador and Newfoundland west to Alberta and south to Minnesota, Wisconsin, New England and New York, then survives in the Appalachians only by climbing, hanging on at 2,500 to 5,000 feet as far south as Virginia and West Virginia. Where it does suit the climate it is superb: a narrow aromatic spire of 50 to 70 feet and only 15 to 25 feet wide, clothed in flat blunt needles 0.6 to 1.2 inches long, dark green above with two chalk-white stomatal bands beneath and twisted at the base so they lie in two horizontal ranks either side of the shoot. Two features are worth learning by hand. The young bark is smooth and grey and studded with RESIN BLISTERS that spray when punctured - no spruce or pine does that - and the erect cones, 2 to 3 inches and dark purple, ripen brown and DISINTEGRATE on the branch in September, dropping their scales and leaving a bare central spike standing on the twig. A fir cone is therefore something you almost never find whole on the ground, which is the single fastest way to tell Abies from Picea. That resin is also the reason this tree has a place in the history of optics: Canada balsam, the oleoresin tapped from those bark blisters, dries to a transparent mass with a refractive index near 1.52, effectively the same as crown glass, so it worked as an invisible glue for cementing lens elements and became the standard mounting medium for microscope slides from about 1830. It was phased out during the Second World War for polyester, epoxy and urethane adhesives, which do not yellow and have far better thermal and solvent resistance. The honest catch list is long: balsam woolly adelgid, spruce budworm (this is the budworm's preferred main host), bark beetles, spider mites in hot spells, cankers, heart rot, root rot, needle rust and twig blight, plus heavy deer browsing, shallow roots that make it wind-firm only in a stand, and a working life measured in decades rather than centuries.